Can you think of how tapetal cells could become bi-nucleate?

The innermost wall layer of microsporangium is known as tapetum. It provides nutrition to the developing microspores. Cells of the tapetum possess dense cytoplasm and generally have more than one nucleus.

Tapetal cells become bi-nucleate by multiple cell fusions of uninuclear cells (cells with a single nucleus). Thus, tapetal cells are syncitium ( cell having more than one nucleus).

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what happen sto the tube nucleus after fertilisation?
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Tapetum is important for the development of pollen grains. The cells are usually bigger and normally have more than one nucleus per cell. As the sporogenous cells undergo mitosis, the nuclei of tapetal cells also divide. Sometimes, this mitosis is not normal due to which many cells of mature tapetum become multinucleate. Sometimes polyploidy and polyteny can also be seen. The unusually large nuclear constitution of the tapetum helps it in providing nutrients and regulatory molecules to the forming pollen grains.
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