In amphibian heart is three chambered.....
why three chambered not four????????? then how his system works without 1 chamber??????

In the course of evolution, amphibians developed 3 chambered heart. One reason for it could be that due to changing environmental conditions and to support the evolved body structure, these organisms developed the multiple chambered heart to adapt successfully with the course of evolution.  

Amphibians have 3 chambered heart; consisting of two atria and one ventricle. The one atria receives oxygenated blood from lungs and other deoxygenated blood from body respectively. The same ventricle first pumps deoxygenated blood to lungs and then oxygenated blood to body with little time gap. There is partial mixing of oxygenated and deoxygenated blood in the ventricle.
They are cold blooded animals and do not need much energy to maintain their body temperature like warm blooded animals (having four chambered heart). Thus, amphibians can allow partial mixing of oxygenated and deoxygenated blood in the ventricle.  
Amphibians also absorb some amount of oxygen through their skin so this supplements the regular respiration. This supplement supply of oxygen through the skin overcomes the absence of two ventricles.  
 

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